Steinhagen, S., Kramár, L. & Toth, G.B. (2022). The unheeded existence of the tubular greens: molecular analyses reveal the distribution of a new Ulva species (Ulvophyceae, Chlorophyta), Ulva capillata sp. nov. in the Atlantic-Baltic Sea transect. J Appl Phycol.

Stedt, K., Toth, G., Davegård, J., Pavia, H., & Steinhagen, S (2022). Determination of nitrogen content in Ulva fenestrata by color image analysis–a rapid and cost-efficient method to estimate nitrogen content in seaweeds. Frontiers in Marine Science, 2510.

Stedt, K., Steinhagen, S., Trigo, J. P.,Kollander, B., Undeland, I., Toth, G. B., Wendin, K. & Pavia, H. (2022). Post-harvest cultivation with seafood process waters improves protein levels of Ulva fenestrata while retaining important food sensory attributes. Front. Mar. Sci. doi: https://10.3389/fmars.2022.991359

Tran, LA.T., Vieira, C., Steinhagen, S., Maggs, C. A., Hiraoka, M., Shimada, S., Nguyen, T. V., De Clercj, O. & Leliaert, F. (2022). An appraisal of Ulva (Ulvophyceae, Chlorophyta) taxonomy. Journal of Applied Phycology.

Steinhagen, S., Larsson, K., Olsson, J., Albers, E., Undeland, I., Pavia, H. & Toth, G. B. (2022). Closed life-cycle aquaculture of sea lettuce (Ulva fenestrata): performance and biochemical profile differ in early developmental stages. Front. Mar. Sci. 9:942679. doi:

Juul, L., Steinhagen, S., Bruhn, A., Jensen, S.K., Undeland, I. & Dalsgaard, T.K. (2022). Combining pressing and alkaline extraction to increase protein yield from Ulva fenestrata biomass. Food and Bioproducts Processing.

van der Loos, L. M., D’hondt, S., Engelen, A. H., Pavia, H., Toth, G. B., Willems, A., Weinberger, F., De Clerck, O., & Steinhagen, S. (2022). Salinity and host drive Ulva-associated bacterial communities across the Atlantic–Baltic Sea gradient. Molecular Ecology, 00, 1– 18.

Steinhagen, S., Enge, S., Cervin, G., Larsson, Edlund, U., Schmidt, A.E., Wahlström, N., Kollander, B., Pavia, H., Undeland, I. & Toth, G. (2022). Harvest time can affect the optimal yield and quality of sea lettuce (Ulva fenestrata) in a sustainable sea-based cultivation. Frontiers in Marine Science. doi:

Stedt, K., Trigo, J.P., Steinhagen, S., Nylund, G.M., Forghani, B., Pavia, H. & Undeland, I. (2022). Cultivation of seaweeds in food production process waters: Evaluation of growth and crude protein content. Algal Research. 63 doi:

Afanasyev, D.F., Steinhagen, S., Abdullin, S.R., & Weinberger, F. (2022). Vegetation of the supralittoral and upper sublittoral zones of the Western German Baltic Sea coast: a phytosociological study. Botanica Marina. 63 doi:


Juul, L., Danielsen, M., Nebel, C., Steinhagen, S., Bruhn, A., Jensen, S.K., Undeland, I. & Dalsgaard T.K. (2021). Ulva fenestrata protein – Comparison of three extraction methods with respect to protein yield and protein quality. Algal Research. doi:

Steinhagen, S., Enge, S., Larsson, K., Nylund, G.M., Albers, E., Pavia, H., Undeland, I. & Toth, G. (2021). Sustainable Large-Scale Aquaculture of the Northern Hemisphere Sea Lettuce, Ulva fenestrata, in an Off-Shore Seafarm. Journal of Marine Science and Engineering. 9(6), 615 doi:

Cai, C., Gu, K., Zhao, H., Steinhagen, S., He, P. & Wichard, T. (2021). Screening and verification of extranuclear genetic markers in green tide algae from the Yellow Sea. PLOS ONE 16(6): e0250968. doi:

Trigo, J.P., Engström, N., Steinhagen, S., Juul, L., Harrysson, H., Toth, G.B., Pavia, H., Scheers, N., & Undeland, I. (2021). In vitro digestibility and Caco-2 cell bioavailability of sea lettuce (Ulva fenestrata) proteins extracted with pH-shift technology. FOOD CHEMISTRY. doi: 10.1016/j.foodchem.2021.129683

Steinhagen, S., Düsedau, L. & Weinberger, F. (2021). DNA barcoding of the German green supralittoral zone indicates the distribution and phenotypic plasticity of Blidingia species and reveals Blidingia cornuta sp. nov.. TAXON. doi: 10.1002/tax.12445


Olsson, J., Raikova, S., Mayers, J.J., Steinhagen, S., Chuck, C.J., Nylund, G.M. & Albers, E. (2020). Effects of geographical location on potentially valuable components in Ulva intestinalis sampled along the Swedish coast. Applied Phycology. 1:1, 80-92, doi: 10.1080/26388081.2020.1827454

Wahlström, N., Steinhagen, S., Toth, G., Pavia, H., Edlund, U. (2020). Ulvan dialdehyde-gelatin hydrogels for removal of heavy metals and methylene blue from aqueous solution. Carbohydrate Polymers. doi: 10.1016/j.carbpol.2020.116841

Toth, G.B., Harrysson, H., Wahlström, N., Olsson, J., Oerbekke, A., Steinhagen, S., Kinnby, A., White, J., Albers, A., Edlund, U., Undeland, I. & Pavia, H. (2020). Effects of irradiance, temperature, nutrients, and pCO2 on the growth and biochemical composition of cultivated Ulva fenestrata. Journal of Applied Phycology. doi: 10.1007/s10811-020-02155-8

Olsson, J., Toth, G.B., Oerbekke, A., Cvijetinovic, S., Wahlström, N., Harrysson, H., Steinhagen, S., Kinnby, A., White, J., Edlund, U., Undeland, I., Pavia, H. & Albers, E. (2020), Cultivation conditions affect the monosaccharide composition in Ulva fenestrata. Journal of Applied Phycology. doi: 10.1007/s10811-020-02138-9


Steinhagen, S., Karez R. & Weinberger, F. (2019). Cryptic, alien, and lost species: Molecular diversity of Ulva sensu lato along the German coasts of the North and Baltic Seas. European Journal of Phycology. doi: 10.1080/09670262.2019.1597925.

Steinhagen, S., Barco, A., Wichard, T. & Weinberger, F. (2019), Conspecificity of the model organism Ulva mutabilis and Ulva compressa (Ulvophyceae, Chlorophyta). Journal of Phycology, 55: 25-36. doi: 10.1111/jpy.12804 

Steinhagen, S., Karez, R. & Weinberger, F. (2019) Molecular analysis of Ulva compressa (Chlorophyta, Ulvales) reveals its morphological plasticity, distribution and potential invasiveness on German North Sea and Baltic Sea coasts. European Journal of Phycology, 54:1, 102-114, doi: 10.1080/09670262.2018.1513167 


Steinhagen, S., Karez, R. & Weinberger, F. (2018). Surveying seaweeds from the Ulvales and Fucales in the world’s most frequently used artificial waterway, the Kiel Canal. Botanica Marina, 62(1), pp. 51-61. doi: 10.1515/bot-2018-0020 

Weinberger, F., Steinhagen, S., Afanasyev, D. &Karez, R. (2018). New records from the southern North Sea and first records from the Baltic Sea of Kornmannia leptoderma. Botanica Marina, 62(1), pp. 63-73. doi: 10.1515/bot-2018-0019 


Afanasyev, D., Kamnev, A., Sushkova, E., Steinhagen, S. (2016). A field guide to Ulvaspecies found in the Black, Azov, Caspian Seas and Eastern Baltic/ Edited by Prof. I.P. Yermakov, Publishing house Pero, Moscow, Russia.

Steinhagen, S. & Weinberger, F. (2016). Ulva und verwandte Grünalgen an Schleswig-Holsteins Küsten – Eine Revision des Arteninventars. Veröffentlichungen des Landesamtes für Landwirtschaft, Umwelt und ländliche Räume Schleswig-Holstein. Vergabenummern: 0608.451410; 0608.451412. 

Dr. Sophie Steinhagen